Godan verb with 'ru' ending, Transitive verb

to clasp; to grasp; to grip; to clutch

to hold (the answer); to have (e.g. the solution); to be the key; to be the reason

to seize (power); to hold (the reins); to dominate; to control

to make (nigirizushi, rice ball, etc.); to form (with one's hands); to press into shape; to mold; to mould